The basic logic for the UpdateTo cell in columns 2 -5 is:
iif(thiscolumn is null and previouscolumn is not null, newvalue, currentvalue)
which will look something like this for field2 (the corresponding entry goes in columns 3,4,5)
iif(isnull(field2) and not isnull(field1), #2010/01/01#, Field2)
For the first column you only need...
iif(isnull(field1), #2010/01/01#,field1)